diff --git a/lib/Cake/Model/Datasource/DboSource.php b/lib/Cake/Model/Datasource/DboSource.php
index ef4bab7e7..5670adab9 100644
--- a/lib/Cake/Model/Datasource/DboSource.php
+++ b/lib/Cake/Model/Datasource/DboSource.php
@@ -2490,16 +2490,16 @@ class DboSource extends DataSource {
 						$count = count($value);
 						if ($count === 1 && !preg_match("/\s+NOT$/", $key)) {
 							$data = $this->_quoteFields($key) . ' = (';
-						} else {
-							$data = $this->_quoteFields($key) . ' IN (';
-						}
-						if ($quoteValues) {
-							if (is_object($model)) {
-								$columnType = $model->getColumnType($key);
+							if ($quoteValues) {
+								if (is_object($model)) {
+									$columnType = $model->getColumnType($key);
+								}
+								$data .= implode(', ', $this->value($value, $columnType));
 							}
-							$data .= implode(', ', $this->value($value, $columnType));
+							$data .= ')';
+						} else {
+							$data = $this->_parseKey($model, $key, $value);
 						}
-						$data .= ')';
 					} else {
 						$ret = $this->conditionKeysToString($value, $quoteValues, $model);
 						if (count($ret) > 1) {
diff --git a/lib/Cake/Test/Case/Model/Datasource/Database/MysqlTest.php b/lib/Cake/Test/Case/Model/Datasource/Database/MysqlTest.php
index 4f3baa0bd..7ae749268 100644
--- a/lib/Cake/Test/Case/Model/Datasource/Database/MysqlTest.php
+++ b/lib/Cake/Test/Case/Model/Datasource/Database/MysqlTest.php
@@ -1960,7 +1960,7 @@ class MysqlTest extends CakeTestCase {
 		$this->assertEquals($expected, $result);
 
 		$result = $this->Dbo->conditions(array('score' => array(2 => 1, 2, 10)));
-		$expected = " WHERE score IN (1, 2, 10)";
+		$expected = " WHERE `score` IN (1, 2, 10)";
 		$this->assertEquals($expected, $result);
 
 		$result = $this->Dbo->conditions("Aro.rght = Aro.lft + 1.1");
@@ -2240,7 +2240,7 @@ class MysqlTest extends CakeTestCase {
 		$this->assertEquals($expected, $result);
 
 		$result = $this->Dbo->conditions(array('score' => array(1, 2, 10)));
-		$expected = " WHERE score IN (1, 2, 10)";
+		$expected = " WHERE `score` IN (1, 2, 10)";
 		$this->assertEquals($expected, $result);
 
 		$result = $this->Dbo->conditions(array('score' => array()));
@@ -2331,7 +2331,7 @@ class MysqlTest extends CakeTestCase {
 			'NOT' => array('Course.id' => null, 'Course.vet' => 'N', 'level_of_education_id' => array(912,999)),
 			'Enrollment.yearcompleted >' => '0')
 		);
-		$this->assertRegExp('/^\s*WHERE\s+\(NOT\s+\(`Course`\.`id` IS NULL\)\s+AND NOT\s+\(`Course`\.`vet`\s+=\s+\'N\'\)\s+AND NOT\s+\(level_of_education_id IN \(912, 999\)\)\)\s+AND\s+`Enrollment`\.`yearcompleted`\s+>\s+\'0\'\s*$/', $result);
+		$this->assertRegExp('/^\s*WHERE\s+\(NOT\s+\(`Course`\.`id` IS NULL\)\s+AND NOT\s+\(`Course`\.`vet`\s+=\s+\'N\'\)\s+AND NOT\s+\(`level_of_education_id` IN \(912, 999\)\)\)\s+AND\s+`Enrollment`\.`yearcompleted`\s+>\s+\'0\'\s*$/', $result);
 
 		$result = $this->Dbo->conditions(array('id <>' => '8'));
 		$this->assertRegExp('/^\s*WHERE\s+`id`\s+<>\s+\'8\'\s*$/', $result);
@@ -2367,7 +2367,7 @@ class MysqlTest extends CakeTestCase {
 
 		$conditions = array('id' => array(2, 5, 6, 9, 12, 45, 78, 43, 76));
 		$result = $this->Dbo->conditions($conditions);
-		$expected = " WHERE id IN (2, 5, 6, 9, 12, 45, 78, 43, 76)";
+		$expected = " WHERE `id` IN (2, 5, 6, 9, 12, 45, 78, 43, 76)";
 		$this->assertEquals($expected, $result);
 
 		$conditions = array('title' => 'user(s)');
diff --git a/lib/Cake/Test/Case/Model/Datasource/DboSourceTest.php b/lib/Cake/Test/Case/Model/Datasource/DboSourceTest.php
index de8cc903f..c38e1d085 100644
--- a/lib/Cake/Test/Case/Model/Datasource/DboSourceTest.php
+++ b/lib/Cake/Test/Case/Model/Datasource/DboSourceTest.php
@@ -1103,4 +1103,73 @@ class DboSourceTest extends CakeTestCase {
 		$this->assertEquals($expected, $result);
 	}
 
+/**
+ * Test conditionKeysToString()
+ *
+ * @return void
+ */
+	public function testConditionKeysToString() {
+		$Article = ClassRegistry::init('Article');
+		$conn = $this->getMock('MockPDO', array('quote'));
+		$db = new DboTestSource;
+		$db->setConnection($conn);
+
+		$conn->expects($this->at(0))
+			->method('quote')
+			->will($this->returnValue('just text'));
+
+		$conditions = array('Article.name' => 'just text');
+		$result = $db->conditionKeysToString($conditions, true, $Article);
+		$expected = "Article.name = just text";
+		$this->assertEquals($expected, $result[0]);
+
+		$conn->expects($this->at(0))
+			->method('quote')
+			->will($this->returnValue('just text'));
+		$conn->expects($this->at(1))
+			->method('quote')
+			->will($this->returnValue('other text'));
+
+		$conditions = array('Article.name' => array('just text', 'other text'));
+		$result = $db->conditionKeysToString($conditions, true, $Article);
+		$expected = "Article.name IN (just text, other text)";
+		$this->assertEquals($expected, $result[0]);
+	}
+
+/**
+ * Test conditionKeysToString() with virtual field
+ *
+ * @return void
+ */
+	public function testConditionKeysToStringVirtualField() {
+		$Article = ClassRegistry::init('Article');
+		$Article->virtualFields = array(
+			'extra' => 'something virtual'
+		);
+		$conn = $this->getMock('MockPDO', array('quote'));
+		$db = new DboTestSource;
+		$db->setConnection($conn);
+
+		$conn->expects($this->at(0))
+			->method('quote')
+			->will($this->returnValue('just text'));
+
+		$conditions = array('Article.extra' => 'just text');
+		$result = $db->conditionKeysToString($conditions, true, $Article);
+		$expected = "(" . $Article->virtualFields['extra'] . ") = just text";
+		$this->assertEquals($expected, $result[0]);
+
+		$conn->expects($this->at(0))
+			->method('quote')
+			->will($this->returnValue('just text'));
+		$conn->expects($this->at(1))
+			->method('quote')
+			->will($this->returnValue('other text'));
+
+		$conditions = array('Article.extra' => array('just text', 'other text'));
+		$result = $db->conditionKeysToString($conditions, true, $Article);
+		$expected = "(" . $Article->virtualFields['extra'] . ") IN (just text, other text)";
+		$this->assertEquals($expected, $result[0]);
+	}
+
 }